「雷獣」か「ジャイアント・ユニコーン」か : 形態比較 : Who's the runner-up?

「雷獣」か「大一角犀」か : Who's the runner-up?　形態比較

 

以下は2018年に投稿した英文記述を、久方ぶりに引っ張り出してきたものですが、今回はスケール調整した二種の形態比較付き。

インドリコテリウム科の超巨大種（パラセラテリウム属種など）を除いた奇蹄類の中で、インドリコテリウム科に次ぐ「二番目（インドリコテリウム科には超巨大種が複数あるゆえ、二番目という表現は正確ではありませんが）」に大きな種類は、大一角犀（エラスモテリウム属最大種＝真正サイ中の最大種）と雷獣（メガセロップス属最大種＝始新世の古代奇蹄類、ブロントテリウム科の最大種）のうち、どちらか？を問いかけた内容で、『プレヒストリック・サファリ㉙』中の和文記述と、対応するものです。

プレヒストリック・サファリ 29　「大型奇蹄類 と 肉食動物②」 '雷獣'(メガセロップス) & ブグティ・ワニ(アストルゴスクス)

以下の復元画で両者の比較ができます。頭骨サイズ、肩高、四肢の長さ・太さなど明確な差異があることが分かると思います。
両者の形態の違いについて、より詳しい記述は上の『プレヒストリック・サファリ㉙』にありますので、ご覧になってください。

概ね同じくらいの推定体重にはなるのでしょうが、平均体重的に、どちらの方が上回っていたと思いますか？

メガセロップス属最大種　'the Thunder Beast'  Megacerops coloradensis

エラスモテリウム属最大種　'the Giant Unicorn Rhino'  Elasmotherium sibiricum/caucasicum

Thunder Beast or Giant Unicorn : Who's the runner-up?

the Saber Panther

 

I've been wondering for sometime, which species was actually the second largest after Paraceratherium transouralicum(and some other super sized forms of indricothere) in the history of perissodactyla.

The middle Pleistocene giant unicorn rhino, Elasmotherium(E. sibiricum or E. caucasicum) is often quoted as one of the largest members of the rhinocerotoidea, perhaps second only to Paraceratherium in size among the entire group. Caucasian and siberian Elasmotherium specimens were indeed humongous, quoted as being over 4.5 m in body length, and over 2 m tall at the shoulders(Zhegallo et al., 2005), thus considerably outsizing the contemporaneous woolly rhinoceros and the extant white rhinoceros. These great ancient rhinos are estimated by some people to have almost matched the woolly mammoth in body mass, perhaps weighing an excess of 4 tons.

However, if there were other perissodactyls that could compete with the giant unicorn for the honorable 'runner-up' placing among the order, that must have been brontotheres, also known as 'thunder beasts'. Being the uncontested largest terrestrial mammals in the Eocene epoch, most known brontotheres were super sized and several species are quoted as reaching unbelievable 2.5 m at the shoulders.

While the claim rings a bit exaggerated, the picture below of a skeleton of Megacerops sp., one of the largest brontotheres, certainly substatiates the claim that they were absolutely super sized perissodactyls. This specimen appears utterly monstrous, incredibly tall and overall more ponderous, especially so at the limbs than does a skeleton of Elasmotherium sp. 

A Megacerops' skull I saw at the Ueno Nature and Science Museum was also huge.

The above mentioned Megacerops specimen is still on display at the American Museum of Natural History, New York, being labeled(erroneously, perhaps) as Brontops robustus.
Is there anyone who has been there and saw the specimen?

Judging from these pictures at hand, this specimen seems easily exceeding 6'11"(2.1 m) in height, around 16 feet(4.9 m) or so in body length, in other words being every inch true to the popularly circulated brontothere measurements. The distal limbs look to be proportionally shorter for such tall stature, more graviportal than does the limbs of Elasmotherium, probably reflecting different ecological adaptation. After all they were not dubbed as 'elephant sized perissodactyls' for nothing.

It's got to be the case that some species of the largest known brontothere genera (Megacerops, Embolotherium or Rhinotitan) were at least comparable to the size of the largest Elasmotherium and accordingly, often quoted body mass estimation of 3 something tons sounds to me a bit too conservative compared to the case of Elasmotherium.

If you have any experience of seeing the skeletons of these behemoths in person, the skeletal measurement data or photos of them taken from similar angles(for better comparison) etc..., please share them with me, that would be awesome!  I'd like to see the image of Elasmotherium skeleton with humans similarly compared as in the Megacerops picture.

Which one do you think must have been the second largest perissodactyl ever?

イラストと本文 by ©the Saber Panther(Jagroar) （All rights reserved）